Dr. rer. nat. Matthias H. Hoffmann



Benítez-Benítez, C., Martín-Bravo, S., Bjorå, C.S., Gebauer, S., Hipp, A.L., Hoffmann, M.H., Luceño, M., Pedersen, T.M., Reznicek, A., Roalson, E., Volkova, P., Yano, O., Spalink, D., Jiménez-Mejías, P. (2021): Geographical vs. ecological diversification patterns in Carex section Phacocystis (Cyperaceae): patterns hidden behind a twisted taxonomy. Journal of Systematics and Evolution 59: 64266. link

Hoffmann, M.H., Gebauer, S., Lühmann, R. (2021): Root anatomy predicts ecological optima in Carex (Cyperaceae) in terms of Ellenberg indicator values. Ecological Indicators 129: 107979. open access

Hoffmann, M.H., Lühmann, R. (2021): The root morphospace of German Carex species and its relation to the phylogeny of the genus. Flora 282: 151882. open access

Roalson, E.H., Jiménez-Mejías, P., Hipp, A.L., Benítez-Benítez, C., Bruederle, L.P., Chung, K.-S., Escudero, M., Ford, B.A., Ford, K., Gebauer, S., Gehrke, B., Hahn, M., Hayat, M.Q., Hoffmann, M.H., Jin, X.-F., Kim, S., Larridon, I., Léveillé-Bourret, É., Lu, Y.-F., Luceño, M., Maguilla, E., Márquez-Corro, J.I., Martín-Bravo, S., Masaki, T., Míguez, M., Naczi, R.F.C., Reznicek, A.A., Spalink, D., Starr, J.R., Uzma, Villaverde, T., Waterway, M.J., Wilson, K.L., Zhang, S. (2021): A framework infrageneric classification of Carex (Cyperaceae) and its organizing principles. Journal of Systematics and Evolution 59: 726762. link


Tkach, N., Schneider, J., Döring, E., Wölk, A., Hochbach, A., Nissen, J., Winterfeld, G., Meyer, S., Gabriel, J., Hoffmann, M.H. and Röser, M. (2020): Phylogenetic lineages and the role of hybridization as driving force of evolution in grass supertribe Poodae. Taxon 69: 234-277. open access

Winterfeld, G., Paule, J., Hoffmann, M. H., Ley, A., Röser, M. (2020): Antagonistic effects of whole-genome duplications and dysploidy on genome sizes in the pantropical monocot family Marantaceae: Consequences in the light of a new molecular phylogeny. Current Plant Biology 24, 100181. open access

Winterfeld, G., Ley, A., Hoffmann, M. H., Paule, J., Röser, M. (2020): Dysploidy and polyploidy trigger strong variation of chromosome numbers in the prayer-plant family (Marantaceae). Plant Systematics and Evolution 306, 36. open access

Villaverde, T., Jiménez-Mejías, P., Luceño, M., Waterway, M. J., Kim, S., Lee, B., Rincón-Barrado, M., Hahn, M., Maguilla, E., Roalson, E. H., Hipp, A. L., Wilson, K. L., Larridon, I., Gebauer, S., Hoffmann, M. H., Simpson, D. A., Naczi, R. F. C., Reznicek, A. A., Ford, B. A., Starr, J. R., Park, J., Escudero, M., Martín-Bravo, S. (2020): A new classification of Carex (Cyperaceae) subgenera supported by a HybSeq backbone phylogenetic tree. Botanical Journal of the Linnean Society 194: 141163.


Hoffmann, M. H. (2019): Text-mining of expert knowledge for the construction of a global habitat space of Micranthes and Saxifraga reveals multiple avenues of arctic biome assembly. International Journal of Plant Sciences 180(3): 240-252.

Hoffmann, M. H. (2019): To the roots of Carex: Unexpected anatomical and functional diversity. Systematic Botany 44(1): 26-31.

Martín-Bravo, S., Jiménez-Mejías, P., Villaverde, T., Escudero, M., Hahn, M., Spalink, D., Roalson, E. H., Hipp, A. L., and the Global Carex Group (Benítez-Benítez, C., Bruederle, L. P., Fitzek, E., Ford, B. A., Ford, K. A., Garner, M., Gebauer, S., Hoffmann, M. H., Jin, X.-F., Larridon, I., Léveillé-Bourret, E., Lu, Y.-F., Luceño, M., Maguilla, E., Márquez-Corro, J. I., Míguez, M., Naczi, R., Reznicek, A. A., Starr, J. R. (2019): A tale of worldwide success: Behind the scenes of Carex (Cyperaceae) biogeography and diversification. Journal of Systematics and Evolution 57: 695-718.


Fläschendräger, A., Hoffmann, M. H. (2018): Anmerkungen zur Höhenverbreitung von Paralaudakia himalayana (Steindachner, 1867). Sauria, Berlin, 40(3): 7577.


Ebersbach, J., Muellner-Riehl, A. N., Michalak, I., Tkach, N., Hoffmann, M. H., Röser, M., Sun, H. and Favre, A. (2017): In and out of the Qinghai-Tibet Plateau: divergence time estimation and historical biogeography of the large arctic-alpine genus Saxifraga L. Journal of Biogeography 44: 900-910. doi:10.1111/jbi.12899

Hoffmann, M. H., Gebauer, S., von Rozycki, T. (2017): Assembly of the arctic flora - highly parallel and recurrent pattern in sedges (Carex). American Journal of Botany 104: 1334-1343. doi:10.3732/ajb.1700133


Global Carex Group1 (2016): Megaphylogenetic specimen-level approaches to the Carex (Cyperaceae) phylogeny using ITS, ETS, and matK sequences: Implications for classification. Systematic Botany 41: 500-518.
1Jiménez-Mejías, P., Hahn, M., Lueders, K., Starr, J. R., Brown, B. H., Chouinard, B. N., Chung, K.-S., Escudero, M., Ford, B. A., Ford, K. A., Gebauer, S., Gehrke, B., Hoffmann, M. H., Jin, X.-F., Jung, J., Kim, S., Luceño, M., Maguilla, E., Martín-Bravo, S., Míguez, M., Molina, A., Naczi, R. F. C., Pender, J. E., Reznicek, A. A., Villaverde, T., Waterway, M. J., Wilson, K. L., Yang, J.-C., Zhang, S., Hipp, A. L., Roalson, E. H.

Global Carex Group2 (2016): Specimens at the center: An informatics workflow and toolkit for specimen-level analysis of public DNA database data. Systematic Botany 41: 529-539.
2Pham, K. K., Hahn, M., Lueders, K., Brown, B. H., Bruederle, L. P., Bruhl, J. J., Chung, K.-S., Derieg, N. J., Escudero, M., Ford, B. A., Gebauer, S., Gehrke, B., Hoffmann, M. H., Hoshino, T., Jiménez-Mejías, P., Jung, J., Kim, S., Luceño, M., Maguilla, E., Martín-Bravo, S., Naczi, R. F. C., Reznicek, A. A., Roalson, E. H., Simpson, D. A., Starr, J. R., Villaverde, T., Waterway, M. J., Wilson, K. L., Yano, O., Zhang, S., Hipp, A. L.

Hochheimer, J., Hoffmann, M. H. (2016): Footprints of the Last Glacial: Past competition may have shaped strongly disjunct distribution ranges in Artemisia rupestris and A. laciniata. Flora 224: 203-210.

Hoffmann, M. H., Gebauer, S. (2016): Quantitative morphological and molecular divergence in replicated and parallel radiations in Carex (Cyperaceae) using Symbolic Data Analysis. Systematic Botany 41: 552-537.


Emadzade, K., Lebmann, M. J., Hoffmann, M. H., Tkach, N., Lone, F. A., Elvira Hörandl, E. (2015): Phylogenetic relationships and evolution of high mountain buttercups (Ranunculus) in North America and Central Asia. Perspectives in Plant Ecology, Evolution and Systematics 17: 131141. link

Gebauer, S., Röser, M., Hoffmann, M. H. (2015): Molecular phylogeny of the species-rich Carex sect. Racemosae (Cyperaceae) based on four nuclear and chloroplast markers. Systematic Botany 40: 433-447.

Global Carex Group1 (2015): Making Carex monophyletic (Cyperaceae, tribe Cariceae): a new broader circumscription. Botanical Journal of the Linnean Society 179: 1-42. open access
1 M. J. Waterway (corresponding author), K. A. Ford, M. Luceño, S. Martín-Bravo, J. R. Starr, K. L. Wilson, O. Yano, S. R. Zhang, E. H. Roalson, W. S. Alverson, L. P. Bruederle, J. J. Bruhl, K.-S. Chung, T. S. Cochrane, M. Escudero, B. A. Ford, S. Gebauer, B. Gehrke, M. Hahn, A. L. Hipp, M. H. Hoffmann, T. Hoshino, P. Jiménez-Mejías, X.-F. Jin, J. Jung, S. Kim, E. Maguilla, T. Masaki, M. Míguez, A. Molina, R. F. C. Naczi, A. A. Reznicek, P. E. Rothrock, D. A. Simpson, D. Spalink, W. W. Thomas, T. Villaverde

Tkach, N., Röser, M., Hoffmann, M. H. (2015): Molecular phylogenetics, character evolution and systematics of the genus Micranthes (Saxifragaceae). Botanical Journal of the Linnean Society 178: 47-66.

Tkach, N., Röser, M., Miehe, G., Muellner-Riehl, A. N., Ebersbach, J., Favre, A., Hoffmann, M. H. (2015): Molecular phylogenetics, morphology and a revised classification of the complex genus Saxifraga (Saxifragaceae). Taxon 64: 1159-1187.


Gebauer, S., Starr, J. R., Hoffmann, M. H. (2014): Parallel and convergent diversification in two northern hemispheric species-rich Carex lineages (Cyperaceae). Organisms Diversity and Evolution 14: 247-258. DOI: 10.1007/s13127-014-0171-9

Tkach, N., Ree, R. H., Kuss, P., Röser, M., Hoffmann, M. H. (2014): High mountain origin, phylogenetics, evolution, and niche conservatism of arctic lineages in the hemiparasitic genus Pedicularis (Orobanchaceae). Molecular Phylogenetics and Evolution 76: 75-92. link


Hoffmann, M. H., Schneider, J., Hase, P., Röser, M. (2013): Rapid and recent world-wide diversification of bluegrasses (Poa, Poaceae) and related genera. PLoS One 8: e60061. doi:10.1371/journal.pone.0060061

Paula-Souza, J. de, Pirani, J. R., Hoffmann, M. H., Röser, M. (2013): Reappraisal of names and lectotype designations in the South American genus Anchietea (Violaceae). Schlechtendalia 25: 6367.


Hoffmann, M. H. (2012): Not across the North Pole: Plant migration in the Arctic. New Phytologist 193: 474-480. doi: 10.1111/j.1469-8137.2011.03924.x


Hoffmann, M. H., Meng, S., Kosachev, P. A., Terechina, T. A., Silanteva, M. M. (2011): Land snail faunas along an environmental gradient in the Altai Mountains (Russia). Journal of Molluscan Studies 77: 76-86.

Schneider, J., Winterfeld, G., Hoffmann, M. H., Röser, M. (2011): Duthieeae, a new tribe of grasses (Poaceae) identified among the early diverging lineages of subfamily Pooideae: molecular phylogenetics, morphological delineation, cytogenetics and biogeography. Systematics and Biodiversity 9(1): 27-44.


Hoffmann, M. H., Paula-Souza, J. de, Fläschendräger, A. Röser, M. (2010): The gynoecium of male Anchietea pyrifolia (Violaceae): preserved structure with a new function. Flora 205: 429-433. doi:10.1016/j.flora.2009.12.001

Hoffmann, M. H., Schmuths, H., Koch, C., Meister, A., Fritsch, R. M. (2010): Comparative analysis of growth, genome size, chromosome numbers and phylogeny of Arabidopsis thaliana and three cooccurring species of the Brassicaceae from Uzbekistan. Journal of Botany, Volume 2010, Article ID 504613, 8 pages, doi:10.1155/2010/504613. link

Hoffmann, M. H., von Hagen, K. B., Hörandl, E. Röser, M., Tkach, N. V. (2010): Sources of the arctic flora: Origins of arctic species in Ranunculus and related genera. International Journal of Plant Sciences. 171(1): 90-106.

Mandle, L., Warren, D. L., Hoffmann, M. H., Peterson, A. T., Schmitt, J., von Wettberg, E. J. (2010): Conclusions about niche expansion in introduced Impatiens walleriana populations depend on method of analysis. PLoS ONE 5(12): e15297. doi:10.1371/journal.pone.0015297


Hoffmann, M. H., Röser, M. (2009): Taxon recruitment of the arctic flora: an analysis of phylogenies. New Phytologist 182: 774-780. pdf - The definitive version is available at http://www.blackwell-synergy.com/loi/nph

Meng, S., Hoffmann, M. H. (2009): Pupilla loessica Lozek 1954 (Gastropoda: Pulmonata: Pupillidae) "A Living Fossil" in Central Asia? Quaternary Science Journal 58: 55-69.

Meng, S., Hoffmann, M. H. (2009): Pupilla seminskii n. sp. (Gastropoda: Pulmonata: Pupilloidea) from the Russian Altay. Mollusca 27(1): 83-86.

Ziermann, J., Ritz, M. S., Hameister, S., Abel, C., Hoffmann, M. H., Neuffer, B., Theißen, G. (2009): Floral visitation and reproductive traits of Stamenoid petals, a naturally occurring floral homeotic variant of Capsella bursa-pastoris (Brassicaceae). Planta 230: 1239-1249. DOI 10.1007/s00425-009-1018-z.


Hoffmann, M. H., Ermakov, N. B. (2008): Biogeographical study of West Siberian hemiboreal forest associations with species range overlay methods. Flora 203(3): 234-242.

Meng, S., Hoffmann, M. H. (2008): Pupilla altaica n. sp. and Pupilla alluvionica n. sp., two new species of Pupillidae (Gastropoda: Pulmonata: Pupilloidea) from the Russian Altay. Mollusca 26(2): 229-234.

Tkach, N. V., Hoffmann, M. H., Röser, M., Korobkov, A. A., von Hagen, K. B. (2008): Parallel evolutionary patterns in multiple lineages of arctic Artemisia L. (Asteraceae). Evolution 62(1): 184-198.

Tkach, N. V., Hoffmann, M. H., Röser, M., von Hagen, K. B. (2008): Temporal patterns of evolution in the Arctic explored in Artemisia L. (Asteraceae) lineages of different age. Plant Ecology & Diversity. 1: 161-169.

Tkach, N. V., Röser, M., Hoffmann, M. H. (2008): Range size variation and diversity distribution in the vascular plant flora of the Eurasian Arctic. Organisms, Diversity and Evolution 8: 251-266.


Hoffmann, M., Ermakov, N. (2007): Modelling plant distribution ranges using climate data. Computational Technologies 12: 178-193 (in English).
Хоффманн, М., Ермаков, Н.Б. (2007): Моделирование ареалов распространения растений с использованием данных по климату. Вычислительные Технологии 12: 178-193 (на английском).


Franke, K., Hoffmann, M., Schmidt, J., Porzel, A., Arnold, N., Wessjohann, L. (2006): 2"-O-Glucosylvitexin, a chemotaxonomic marker for the genus Cryptocoryne (Araceae). Biochemical Systematics and Ecology 34: 546-548.

Harpke, D., Peterson, A., Hoffmann, M. H., Röser, M. (2006): Phylogenetic evaluation of chloroplast trnL-trnF DNA sequence variation in the genus Mammillaria (Cactaceae). Schlechtendalia 14: 7-16.

Schmid, K. J., Törjék, O., Meyer, R., Schmuths, H., Hoffmann, M. H., Altmann, T. (2006): Evidence for a large-scale population structure of Arabidopsis thaliana from genome-wide single nucleotide polymorphism markers. Theoretical and Applied Genetics 112: 1104-1114.

Schmuths, H., Bachmann, K., Weber W. E., Horres, R., Hoffmann, M. H. (2006): Effects of preconditioning and temperature during germination of 73 natural accessions of Arabidopsis thaliana. Annals of Botany 97: 623-634.

Хоффман М. (2006): Два подхода к моделированию климатических ареалов видов растений. Шумный В.К., Шокин Ю.И., Колчанов Н.А., Федотов А.М. (Ред.): Биоразнообразие и динамика экосистем: информационные технологии и моделирование. Новосибирск, Издательство Сибирского Отделения РАН, 2006, с. 289-301.
Hoffmann M. (2006): Two approaches of climatic range modelling of plant species. In: Shumny, V. K., Shokin, Yu. I., Kolchanov, N. A., Fedotov, A. M. (eds.): Biodiversity and Dynamic of Ecosystems: Computational Approaches and Modelling. Novosibirsk, Publishing House of the Siberian Branch of the Russian Academy of Sciences. pp. 289-301 (In Russian).


Hoffmann, M. H. (2005): Evolution of the realized climatic niche in the genus Arabidopsis (Brassicaceae). Evolution 59(7): 1425-1436.

Hoffmann, M. H., Tomiuk, J., Schmuths, H., Koch, C., Bachmann, K. (2005): Phenological and morphological responses to different temperature treatments differ among a world-wide sample of accessions of Arabidopsis thaliana. Acta Oecologica 28: 181-187.

Tan, Y. -Y., Xu, H. -H., Tao, W. -J., Hoffmann, M. H., Wang, X. -F., Lu, Y. -T. (2005): Transgenic GFP as a molecular marker for approaches to quantify pollination mechanism and gene flow in Arabidopsis thaliana. Plant Biology 7: 405-410.


Döring, E., Hoffmann, M. H. (2004): Distribution analysis of monophagous butterflies (Lepidoptera) and their host plants in the western Palaearctic. Schlechtendalia 12: 85-93.

Schmuths, H., Hoffmann, M. H., Bachmann, K. (2004): Geographic distribution and recombination of genomic fragments on the short arm of chromosome 2 of Arabidopsis thaliana. Plant Biology 6: 128-139.


Hoffmann, M. H., Bremer, M., Schneider, K., Burger, F., Stolle, E., Moritz, G. (2003): Flower visitors in a natural population of Arabidopsis thaliana. Plant Biology 5: 491-494.

Hoffmann, M. H., Glaß, A. S., Tomiuk, J., Schmuths, H., Fritsch, R. M., Bachmann, K. (2003): Analysis of molecular data of Arabidopsis thaliana (L.) Heynh. (Brassicaceae) with Geographical Information Systems (GIS). Molecular Ecology 12, 1007-1019.


Hoffmann, M. H. (2002) Biogeography of Arabidopsis thaliana (L.) Heynh. (Brassicaceae). Journal of Biogeography 29: 125-134.

Hoffmann, M. H., Welk, E., Zapp, M. (2002): Climatic mean diagrams: A technique for assessing the climatic differentiation of species. Plant Systematics and Evolution 231: 191-202.

Welk, E., Schubert, K., Hoffmann, M. H. (2002): Present and potential distribution of invasive garlic mustard (Alliaria petiolata) in North America. Diversity and Distributions 8: 219-233.


Hoffmann, M. H. (2001) The distribution of Senecio vulgaris: capacity of climatic range models for predicting adventitious ranges. Flora 196, 395-403.

Hoffmann, M. H. (2001): Klimatische Modellierung des Areals von Scleranthus annuus. In: Brandes, D. (Hrsg.): Adventivpflanzen. Beiträge zur Biologie, Vorkommen und Ausbreitungsdynamik von Archäophyten und Neophyten in Mitteleuropa. Braunschweiger Geobotanische Arbeiten 8: 169-178.

Hoffmann, M. H., Schmuths, H. (2001): Arabidopsis thaliana as a tool for biosystematics: studies in molecular phylogeography. IOPB Newsletters 33, 17-21.

Hoffmann, M. H., Telyatnikov, M. Yu., Ermakov, N. (2001): Phytogeographical analysis of plant communities along an altitudinal transect through the Kuraiskaya basin (Altai, Russia). Phytocoenologia 31, 401-426.


Hoffmann, M. H. (2000) Biogeography and climatic differentiation of two annual species of Teesdalia R. Br. (Brassicaceae). Journal of Biogeography 27, 989-999.

Hoffmann, M. H. (2000): Type specimens of the Brassicaceae in the Herbarium of the Martin-Luther-University Halle-Wittenberg (HAL). Schlechtendalia 4, 35-40.

Kesting, S., Hoffmann, M. H. (2000): Type specimens of the Apocynaceae, Asclepiadaceae, Gentianaceae, Loganiaceae, and Menyanthaceae in the Herbarium of the Martin-Luther-University Halle-Wittenberg (HAL). Schlechtendalia 5, 11-17.

Welk, E., Hoffmann, M. H. (2000): Bewertung der Schutzrelevanz von Gefäßpflanzen in Deutschland aus arealkundlicher Sicht. Schriftenreihe für Landschaftspflege Naturschutz 68, 89-109.


Hoffmann, M. H. (1999): The phylogeny of Actaea (Ranunculaceae): a biogeographical approach. Plant Systematics and Evolution 216, 251-263.

Hoffmann, M. H. (1999): Biogeographical and evolutionary patterns in the genus Caltha L. (Ranunculaceae). Botanische Jahrbücher Systematik 121, 403-421.

Hoffmann, M. H. (1999): Geographische Musterbildungen bei Sippendifferenzierungen. Diss. Martin-Luther-Universität Halle-Wittenberg.

Hoffmann, M. H., Welk, E. (1999): A method for the estimation of the global population sizes of plant species - the Area-abundance-index. Global Ecology and Biogeography 8, 39-46.


Hoffmann, M. H. (1998): Ecogeographical differentiation patterns in Adonis sect. Consiligo (Ranunculaceae). Plant Systematics and Evolution 211, 43-56.

Hoffmann, M. H. (1998): Type specimens of the Ranunculaceae in the Herbarium of the Martin-Luther-University Halle-Wittenberg (HAL). Schlechtendalia 1, 19-22.

Hoffmann, M. H., Litt, T., Jäger, E. J. (1998): Ecology and climate of the early Weichselian flora from Gröbern (Germany). Review of Palaeobotany and Palynology. 102, 259-276.

Welk, E., Hoffmann, M. H. (1998): Chorologische Datenbanken - grundlegende Voraussetzung zur objektiven Evaluierung der Schutzrelevanz von Gefäßpflanzen. Zeitschrift für Ökologie und Naturschutz 7, 155-168.


Jäger, E. J., Hoffmann, M. H. (1997): Schutzwürdigkeit von Gefäßpflanzen aus der Sicht der Gesamtareale. Zeitschrift für Ökologie und Naturschutz 6, 225-232.

Hoffmann, M. H. (1996): Die in Zentraleuropa verwilderten und kultivierten nordamerikanischen Astern. Feddes Repertorium 107(3-4), 163-188.

Hoffmann, M. H. (1995): Die in Europa eingeschleppten und verwilderten nordamerikanischen Astern. Information zur Floristischen Kartierung von Thüringen 9, 17-20.

M. H. Hoffmanns homepage